1. Department of Biological Sciences, Graduate School of Science, the University of Tokyo, Tokyo, Japan
2. Department of Anatomy, St. Marianna University School of Medicine, Kanagawa, Japan
3. Laboratory of Physiology, Atmosphere and Ocean Research Institute, the University of Tokyo, Chiba, Japan
4. Department of Aquatic Bioscience, Graduate School of Agricultural and Life Sciences, the University of Tokyo, Tokyo, Japan