1. College of Veterinary Medicine, Yangzhou University , Yangzhou 225009, Jiangsu, People's Republic of China
2. State Key Laboratory of Farm Animal Biotech Breeding, College of Biological Sciences, China Agricultural University , Beijing 10021, People's Republic of China
3. Joint International Research Laboratory of Agriculture and Agri-Product Safety, The Ministry of Education of China, Yangzhou University , Yangzhou 225009, Jiangsu, People's Republic of China
4. Jiangsu Co-innovation Center for Prevention and Control of Important Animal Infectious Diseases and Zoonoses , Yangzhou 225009, Jiangsu, People's Republic of China