1. Department of Aquatic Bioscience, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo, Tokyo, Japan
2. Laboratory for Systems Molecular Ethology, RIKEN Center for Brain Science, Wako, Saitama, Japan
3. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo, Tokyo, Japan
4. Department of Anatomy and Neurobiology, Graduate School of Medicine, Nippon Medical School, Bunkyo, Tokyo, Japan