1. Department of Biological Sciences, Graduate School of Science, University of Tokyo, Tokyo, Japan
2. Division of Biology and Biological Engineering, California Institute of Technology, Pasadena, California
3. Department of Anatomy, St. Marianna University School of Medicine, Kanagawa, Japan
4. Graduate School of Bioagricultural Sciences, Nagoya University, Aichi Japan
5. Department of Life Science, International Christian University, Tokyo, Japan