1. Departments of Animal Sciences and Applied Biological Chemistry, Graduate School of Agriculture and Life Sciences, The University of Tokyo, Tokyo, Japan
2. Faculty of Natural System, Institute of Science and Engineering, Kanazawa University, Noto Marine Laboratory, Noto, Japan
3. Juntendo University Graduate School of Medicine, Tokyo, Japan
4. Department of Molecular, Cellular and Developmental Biology, University of Michigan, Ann Arbor, Michigan