1. National Park Service, Glacier Bay National Park and Preserve, PO Box 140, Gustavus, AK 99826 USA; School of Environmental and Forestry Sciences, University of Washington, Seattle, WA 98195 USA;
2. National Park Service, Glacier Bay National Park and Preserve, P.O. Box 140, Gustavus, AK 99826 USA
3. School of Environmental and Forestry Sciences, University of Washington, Seattle, WA 98195 USA