1. Laboratory of Plant Genetics and Breeding Science, Department of Agriculture, Faculty of Agriculture, Tokyo University of Agriculture, 1737 Funako, Atsugi City, Kanagawa Prefecture 243-0034, Japan
2. Laboratory of Plant Chromosome and Gene Stock, Graduate School of Science, Hiroshima University, Higashi-Hiroshima City, Hiroshima 739-8526, Japan
3. South Siberian Botanical Garden and Department of Plant Systematics, Altai State University, Barnaul 656099, Russia
4. Research Institute of Evolutionary Biology, 2-4-28 Kamiyouga, Setagaya-Ku, Tokyo, Japan