1. Department of Orthopaedic Surgery Perelman School of Medicine, University of Pennsylvania Philadelphia PA
2. Department of Orthopaedic Surgery The First Hospital of China Medical University Shenyang China
3. Department of Basic & Translational Sciences School of Dental Medicine, University of Pennsylvania Philadelphia PA
4. Department of Cell and Developmental Biology Perelman School of Medicine, University of Pennsylvania Philadelphia PA USA
5. Penn Institute for Regenerative Medicine, Musculoskeletal Program, Perelman School of Medicine, University of Pennsylvania Philadelphia PA USA