1. Animal Postcranial Evolution Laboratory, Skeletal Biology Research Centre, School of Anthropology and Conservation; University of Kent; Canterbury UK
2. Durrell Institute of Conservation and Ecology, School of Anthropology and Conservation; University of Kent; Canterbury UK
3. School of Mathematics, Statistics and Actuarial Science; University of Kent; UK
4. Department of Human Evolution; Max Planck Institute for Evolutionary Anthropology; Leipzig Germany