1. Research Unit of Biology and Applied Ecology, Department of Animal Biology, Faculty of SciencesUniversity of Dschang Dschang Cameroon
2. Research Unit of Soil Analysis and Environmental Chemistry, Department of Soil Sciences, Faculty of Agronomy and Agricultural SciencesUniversity of Dschang Dschang Cameroon
3. Laboratory of General Biology, Department of Biology and Animal Physiology, Faculty of SciencesUniversity of Yaounde I Yaounde Cameroon