1. School of Biosciences The University of Melbourne Parkville Victoria 3010 Australia
2. Unité de Mathématiques et Informatique Appliquées (MIAT) Toulouse INRA Auzeville BP 52627 31326 Cedex France
3. RSPB Centre for Conservation Science, RSPB The Lodge Sandy Bedfordshire SG19 2DL United Kingdom
4. United States Geological Survey Patuxent Wildlife Research Centre 12100 Beech Forest Road Laurel Maryland 20708 USA
5. Centre for Biodiversity and Conservation Science School of Geography, Planning and Environmental Management University of Queensland St Lucia Queensland 4069 Australia