1. Environmental Sciences Division Oak Ridge National Laboratory Oak Ridge Tennessee 37831 USA
2. School of Environment and Natural Resources Ohio State University Columbus Ohio 43210 USA
3. Department of Ecology and Evolutionary Biology University of Tennessee Dabney Hall, 1416 Circle Drive Knoxville Tennessee 37996 USA
4. Tvärminne Zoological Station University of Helsinki J.A. Palménin tie 260 Hanko 10900 Finland
5. Horn Point Laboratory University of Maryland Center for Environmental Science 2020 Horns Point Road Cambridge Maryland 21613 USA