1. Laboratory of Plant Biodiversity and Dynamics of Ecosystems in Arid Environment Faculty of Sciences of Sfax University of Sfax Soukra Road Sfax 3038 Tunisia
2. Research Unit of Macromolecular Biochemistry and Genetic (BMG) Department of Life Sciences Faculty of Sciences of Gafsa University of Gafsa Campus Universitaire Sidi Ahmed Zarroug Gafsa 2112 Tunisia
3. Laboratory of Environmental Bioprocesses Center of Biotechnology of Sfax, AUF (PER‐LBP) Route de Sidi Mansour Sfax 1177 Tunisia