1. Department of Biomedicine; Graduate School of Medical Sciences, Kyushu University; Fukuoka Japan
2. Center of Biomedical Research, Research Center for Human Disease Modeling, Graduate School of Medical Sciences, Kyushu University; Fukuoka Japan
3. Joint Faculty of Veterinary Medicine, Transboundary Animal Diseases Research Center, Kagoshima University; Kagoshima Japan
4. Department of Ophthalmology; Graduate School of Medical Sciences, Kyushu University; Fukuoka Japan
5. Shiga Prefectural Institute of Public Health; Shiga Japan
6. Department of Animal Medical Sciences; Faculty of Life Sciences, Kyoto Sangyo University; Kyoto Japan