1. Academic Unit of Ophthalmology, School of Clinical Sciences; University of Bristol; Bristol UK
2. University College London-Institute of Ophthalmology; London UK
3. Department of Clinical Medicine, School of Medicine; Trinity College Dublin; Dublin Ireland
4. Smurfit Institute of Genetics; Trinity College Dublin; Dublin Ireland
5. National Institute for Health Research (NIHR) Biomedical Research Centre at Moorfields Eye Hospital and University College London Institute of Ophthalmology; London UK