1. State Key Laboratory of South China Sea Marine Resource Utilization; Hainan University; Haikou P. R. China
2. Institute of Tropical Agriculture and Forestry; Hainan University; Haikou P. R. China
3. Department of Molecular Biosciences and Bioengineering; University of Hawaii at Manoa; Honolulu USA
4. Ministry of Eduction Key Laboratory of Tropical Island Ecology; Hainan Normal University; Haikou P. R. China