1. Laboratory of Plant Systematics and Evolutionary Biology College of Life Sciences, Wuhan University Wuhan 430072 China
2. Ecology and Environment Monitoring and Scientific Research Center, Yangtze Basin Ecology and Environment Administration, Ministry of Ecology and Environment of the People's Republic of China Wuhan 430010 China
3. State Key Laboratory of Hybrid Rice, Laboratory of Plant Systematics and Evolutionary Biology College of Life Sciences, Wuhan University Wuhan 430072 China