1. Centre for Ecology & Conservation; College of Life and Environmental Sciences; University of Exeter; Penryn Campus; Penryn TR10 9EZ U.K
2. Conservation and Fisheries Department; Ministry of Natural Resources and Labour; PO Box 3323 Road Town Tortola British Virgin Islands
3. The Settlement; Anegada British Virgin Islands
4. Environment and Sustainability Institute; University of Exeter; Penryn Campus; Penryn Cornwall TR10 9EZ U.K