1. Department of Biology University of Maryland College Park Maryland 20744 USA
2. Department of Integrative Biology and Ecology, Evolutionary Biology, and Behavior Program Michigan State University East Lansing Michigan 48824 USA
3. U.S. Geological Survey Patuxent Wildlife Research Center SO Conte Anadromous Fish Research Lab Turners Falls Massachusetts 01376 USA
4. U.S. Geological Survey Patuxent Wildlife Research Center Laurel Maryland 20708–4017 USA
5. Department of Ecology & Evolutionary Biology Cornell University Ithaca New York 14583 USA