1. Key Laboratory of Engineering Biology for Low-carbon Manufacturing Institute of Industrial Biotechnology Chinese Academy of Science 32 West 7th Avenue Tianjin 300308 P. R. China
2. School of Chemical Engineering and Technology Xi'an Jiaotong University No.28, Xianning West Road Xi'an Shaanxi 710049 P. R. China
3. Department of Biotechnology Delft University of Technology van der Maasweg 9 2629HZ Delft The Netherlands