1. Groningen Institute for Evolutionary Life Sciences; University of Groningen; P.O. Box 11103 9700CC Groningen The Netherlands
2. Department of Ecological Science; Vrije Universiteit; De Boelelaan 1085 1081 HV Amsterdam The Netherlands
3. Centre for Biodiversity Theory and Modeling, Theoretical and Experimental Ecology Station; CNRS and Paul Sabatier University; 09200 Moulis France