1. Faculty of Resource Science and Technology Universiti Malaysia Sarawak 94300 Kota Samarahan Sarawak Malaysia
2. Penang Botanic Gardens Department Kompleks Pentadbiran, Bangunan Pavilion, Jalan Kebun Bunga 10350 Pulau Pinang Malaysia
3. Research Associate Harvard University Herbaria 22 Divinity Avenue Cambridge MA 02138 United States
4. Ludwig‐Maximilians‐Universität München, Department Biologie I Systematische Botanik und Mykologie Menzinger Straße 67 80638 München Germany