1. Charles Perkins Centre The University of Sydney Sydney New South Wales Australia
2. School of Mathematics and Statistics The University of Sydney Sydney New South Wales Australia
3. School of Life and Environmental Sciences Faculty of Veterinary Science The University of Sydney Sydney New South Wales Australia
4. San Diego Zoo Global San Diego California 92112 USA
5. Department of Ecology and Evolutionary Biology University of Toronto Toronto Ontario M5S 3B2 Canada
6. Evolution and Ecology Research Centre, and School of Biological, Earth and Environmental Sciences University of New South Wales Sydney New South Wales Australia
7. Department of Chemistry and Biology Ryerson University Toronto Ontario M5B 2K3 Canada
8. BECO do Departamento de Zoologia Instituto de Biociências Universidade de São Paulo Rua do Matão, trav. 14, no. 321 São Paulo São Paulo 05508‐090 Brazil