1. Department of Plant Sciences; University of Oxford; South Parks Road Oxford OX1 3RB U.K.
2. Royal Botanic Garden Edinburgh; 20A Inverleith Row Edinburgh EH5 3LR U.K.
3. Herbario MOL, Departamento de Manejo Forestal; Facultad de Ciencias Forestales; Universidad Nacional Agraria La Molina; Apartado 456 La Molina, Lima Peru
4. Embrapa Recursos Genéticos e Biotechnologia, PqEB; Caixa Postal 02372 Brasilia-DF 70770-917 Brazil
5. Institute for Systematic Botany; University of Zurich; Zollikerstrasse 107 8008 Zurich Switzerland