1. Department of Evolutionary Ecology; Estación Biológica de Doñana-CSIC; Av. Américo Vespucio 41092 Seville Spain
2. Department of Integrative Sciences; University of Huelva; Campus Universitario El Carmen Av. Andalucía 21071 Huelva Spain
3. Department of Conservation Biology; Estación Biológica de Doñana-CSIC; Av. Américo Vespucio 41092 Seville Spain
4. Centre for Applied Ecology “Prof. Baeta Neves”/InBIO; Institute Superior of Agronomy; University of Lisbon; Tapada da Ajuda 1349-017 Lisboa Portugal