1. Departments of Molecular GeneticsBiochemistry and ChemistryUniversity of Toronto Toronto Ontario M5S 1A8 Canada
2. TIFR Centre for Interdisciplinary SciencesTata Institute of Fundamental Research Hyderabad Telangana 500107 India
3. Program in Molecular MedicineHospital for Sick Children 555 University Avenue Toronto Ontario M5G 1X8 Canada