1. Graduate School of Pharmaceutical Sciences; The University of Tokyo; 7-3-1 Hongo, Bunkyo-ku Tokyo 113-0033 Japan
2. Graduate School of Engineering; The University of Tokyo; JST-ACCEL; 7-3-1 Hongo, Bunkyo-ku Tokyo 113-8656 Japan
3. RIKEN Center for Life Science Technologies, Yokohama; Kanagawa 230-0045 Japan