1. Graduate School of Medicine The University of Tokyo 7-3-1 Hongo, Bunkyo-ku Tokyo 113-0033 Japan
2. Graduate School of Pharmaceutical Sciences The University of Tokyo 7-3-1 Hongo, Bunkyo-ku Tokyo 113-0033 Japan
3. PRESTO Japan Science and Technology Agency 4-1-8 Honcho, Kawaguchi Saitama 332-0012 Japan
4. School of Pharmaceutical Sciences University of Geneva University of Lausanne 1 Rue Michel Servet 1211 Geneva Switzerland
5. CREST Japan Agency for Medical Research and Development (AMED) 1-7-1 Otemachi, Chiyoda-ku Tokyo 100-0004 Japan