1. Graduate School for Life and Environmental Sciences; Kyoto Prefectural University; 1-5 Shimogamo Hangi-cho, Sakyo-ku Kyoto 606-8522 Japan
2. Faculty of Advanced Life Science; Frontier Research Center for Post-Genome Science and Technology; Hokkaido University; Kita, 21 Nishi 11 Sapporo 001-0021 Japan
3. Department of Applied Biological Science; Faculty of Science and Technology; Tokyo University of Science; 2641 Yamazaki, Noda Chiba 278-8510 Japan