1. Graduate School of Medicine; The University of Tokyo; 7-3-1, Hongo Bunkyo-ku Tokyo 113-0033 Japan
2. PRESTO, Japan, Science and Technology Agency; 4-1-8 Honcho Kawaguchi Saitama 332-0012 Japan
3. Graduate School of Pharmaceutical Sciences; The University of Tokyo; 7-3-1 Hongo Bunkyo-ku Tokyo 113-0033 Japan
4. Division of Molecular Neurobiology; National Institute for Basic Biology; 5-1 Higashiyama Myodaiji-cho, Okazaki Aichi 444-8787 Japan
5. School of Life Science; The Graduate University for Advanced Studies; 5-1 Higashiyama Myodaiji-cho, Okazaki Aichi 444-8787 Japan
6. Life Science Center, Tsukuba Advanced Research Alliance; Tsukuba Ibaraki 305-8577 Japan
7. Graduate School of Life and Environmental Sciences; University of Tsukuba; Tsukuba Ibaraki, 305-8577 Japan
8. CREST, Japan, Agency for Medical Research and Development; 1-7-1 Otemachi Chiyoda-ku Tokyo 100-0004 Japan