1. Faculty of Biosciences and Aquaculture; University of Nordland; 8049; Bodø; Norway
2. School of Botany; University of Melbourne; Victoria; 3010; Australia
3. Shoals Marine Laboratory; Cornell University; Portsmouth; New Hampshire; 03801
4. Marine Benthic Ecology and Evolution Group; Centre for Ecological and Evolutionary Studies; University of Groningen; Nijenborgh 7; 9747 AG; Groningen; The Netherlands