1. School of Biological Sciences Coastal & Marine Ecosystems Group University of Sydney Sydney New South Wales 2000 Australia
2. CSIRO Oceans and Atmosphere – Coasts GPO Box 1538 Hobart Tasmania 7001 Australia
3. CSIRO National Research and Collections Australia GPO Box 1538 Hobart Tasmania 7001 Australia
4. School of Biological, Earth and Environmental Sciences Evolution & Ecology Research Centre UNSW Sydney New South Wales 2052 Australia