1. Faculty of Biology Moscow State Lomonosov University Leninskie Gory 1‐12 Moscow119234Russia
2. Plant Ecology & Nature Conservation Group Department of Environmental Science Wageningen University & Research Centre Wageningen The Netherlands
3. Systems Ecology Department of Ecological Science Faculty of Science Vrije Universiteit De Boelelaan 1085 Amsterdam1081 HVThe Netherlands
4. Teberda State Reserve Badukskii 1Karachaevo‐Cherkessian Republic Teberda369210Russia
5. Timiryazev Institute of Plant Physiology RAS Botanicheskaya ul. 35 Moscow127276Russia
6. Institute of Ecology and Earth Sciences University of Tartu Lai 40 Tartu51005Estonia
7. Faculty of Soil Science Moscow State Lomonosov University Leninskie Gory 1‐12 Moscow119234Russia