1. Univ. Lille, CNRS Univ. Littoral Côte d’Opale UMR 8187 – LOG – Laboratoire d’Océanologie et de Géosciences F‐59000Lille France
2. Department of Marine Resources and Energy Tokyo University of Marine Science and Technology 4‐5‐7 Konan, Minato‐ku Tokyo108‐8477Japan
3. Department of Zoology and Entomology Rhodes University Grahamstown6140South Africa
4. CCMAR–Centro de Ciencias do Mar CIMAR Laboratório Associado Universidade do Algarve Campus de Gambelas Faro8005‐139Portugal