1. USDA Forest ServicePacific Northwest Research Station Juneau AK 99801 USA
2. Department of Biological SciencesUniversity of Notre Dame Notre Dame IN 46556 USA
3. Biology DepartmentNorthern Michigan University Marquette MI 49855 USA
4. US Army Corps of EngineersEngineer Research and Development Center Vicksburg MS 39180 USA
5. Institute of Ecology and EvolutionUniversity of Oregon Eugene OR 97403 USA