1. School of Public Health Zhejiang University Hangzhou China
2. Centre for Global Health, Usher Institute University of Edinburgh Edinburgh UK
3. Colon Cancer Genetics Group, Cancer Research UK Edinburgh Centre and Medical Research Council Human Genetics Unit, Medical Research Council Institute of Genetics and Molecular Medicine University of Edinburgh Edinburgh UK
4. Cancer Research UK Edinburgh Centre Medical Research Council Institute of Genetics and Molecular Medicine, University of Edinburgh Edinburgh UK
5. Danish Institute for Advanced Study (DIAS), Department of Public Health, University of Southern Denmark Odense Denmark
6. Centre for Population Health Sciences Usher Institute, University of Edinburgh Edinburgh UK
7. Division of Genetics and Epidemiology The Institute of Cancer Research London UK