1. Centre for Invasion Biology, Department of Zoology and Entomology; University of Pretoria; Private Bag X 20 Hatfield 0028 South Africa
2. South African Institute for Aquatic Biodiversity; Grahamstown 6140 South Africa
3. Department of Life Sciences; Imperial College London, Silwood Park Campus; Buckhurst Road Ascot Berkshire SL5 7PY
4. Centre for Invasion Biology, School of Animal, Plant and Environmental Sciences; University of the Witwatersrand; Private Bag 3, Wits Johannesburg 2050 South Africa
5. Centre for Invasion Biology; South African Institute for Aquatic Biodiversity; Grahamstown 6140 South Africa