1. Center for Limnology University of Wisconsin‐Madison 680 N. Park Street Madison Wisconsin 53706 USA
2. Institute of Low Temperature Science Hokkaido University Kita‐19, Nishi‐8, Kita‐ku Sapporo 060‐0819 Japan
3. Japan Agency for Marine‐Earth Science and Technology 2‐15 Natsushima‐cho Yokosuka 237‐0061 Japan