1. University of Johannesburg; Department of Botany and Plant Biotechnology; P.O. Box 524, Auckland Park 2006, Johannesburg South Africa
2. South African National Biodiversity Institute; Compton Herbarium; Private Bag X7, Claremont 7735 South Africa
3. Universidad Nacional del Comahue; INIBIOMA, CONICET -; Bariloche, Río Negro 8400 Argentina
4. Harbin Normal University; Department of Biology; Hexing Road 50, Harbin 150080 People's Republic of China
5. University of Illinois at Urbana-Champaign; Department of Plant Biology; Urbana, Illinois 61801 U.S.A