1. University of Pretoria; Schweickerdt Herbarium, Department of Plant Science; Pretoria, 0002 South Africa
2. Brooklyn Botanic Garden; 1000 Washington Avenue, Brooklyn, New York 11225 U.S.A
3. South African National Biodiversity Institute, / Acocks Chair, Schweickerdt Herbarium, Department of Plant Science, University of Pretoria, Pretoria, 0002 South Africa; Office of the Chief Director: Research and Scientific Services; University of Pretoria; Acocks Chair, Schweickerdt Herbarium, Department of Plant Science; Private Bag X101, Pretoria, 0001 South Africa , Pretoria, 0002 South Africa