1. Department of Evolution, Ecology and Organismal Biology; The Ohio State University; 300 Aronoff Laboratory, 318 W. 12th Avenue, Columbus, Ohio 43210 U.S.A
2. Department of Biological Sciences; Sam Houston State University; Huntsville, Texas 77340 U.S.A
3. Rancho Santa Ana Botanic Garden; 1500 North College Avenue, Claremont, California 91711 U.S.A