1. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province Chengdu Institute of Biology, Chinese Academy of Sciences 610041 Chengdu China
2. State Key Laboratory of Vegetation and Environmental Change Institute of Botany, Chinese Academy of Sciences 100093 Beijing China
3. University of Chinese Academy of Sciences No. 19A Yuquan Road 100049 Beijing China