1. State Key Laboratory of Forest and Soil Ecology, Institute of Applied Ecology, Chinese Academy of Sciences, 72 Wenhua Road, Shenyang, 110164, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Department of Natural Resource Sciences, Macdonald Campus, McGill University, 21, 111 Lakeshore Road, Ste‐Anne‐de‐Bellevue, QC, Canada H9X 3V9