1. School of Biology University of St Andrews St Andrews KY16 9TH United Kingdom
2. Organismal and Evolutionary Biology Research Programme University of Helsinki PO Box 65 FI‐00014 Finland
3. Centre of Excellence in Biological Interactions
4. Ecology and Genetics Research Unit University of Oulu POB 3000 FI‐90014 Finland
5. Department of Biological and Environmental Science University of Jyvaskyla PO Box 35 FI‐40014 Finland