1. Laboratory of Microbial Technology, Division of Systems Bioengineering, Department of Bioscience and Biotechnology, Faculty of AgricultureGraduate School, Kyushu University, Motooka Nishi‐ku Fukuoka Japan
2. Botany and Microbiology Department, Faculty of Science for BoysAl‐Azhar University, Nasr city Cairo Egypt
3. Laboratory of Soil and Environmental Microbiology, Division of Systems Bioengineering, Department of Bioscience and Biotechnology, Faculty of AgricultureGraduate School, Kyushu University, Motooka Nishi‐ku Fukuoka Japan
4. Laboratory of Microbial Environmental Protection, Tropical Microbiology UnitCenter for International Education and Research of Agriculture, Faculty of Agriculture, Kyushu University 744, Motooka Nishi‐ku Fukuoka Japan