1. The Mechanobiology Institute, National University of Singapore; 5A Engineering Drive 1 Singapore 117411 Singapore
2. Engineering Department; Cambridge University; Trumpington Street Cambridge CB2 1PZ UK
3. Department of Molecular Cell Biology; Weizmann Institute of Science; Rehovot 76100 Israel
4. Laboratoire Matière et Systèmes Complexes; UMR 7057 CNRS and Université Paris Diderot; 10 Rue Alice Domon et Léonie Duquet Paris 75013 France