1. CGBTC-China-Germany Brewing Technical Centre, Shandong Provincial Key Laboratory of Microbial Engineering; Shandong University of Technology; Jinan Shandong 250353 People's Republic of China
2. Key Laboratory of Industrial Fermentation Microbiology (Tianjin University of Science and Technology); Ministry of Education; Tianjin 300457 People's Republic of China
3. School of Life Sciences; Heriot-Watt University; Edinburgh EH14 4AS UK
4. ICBD-International Centre for Brewing and Distilling; Heriot-Watt University; Edinburgh EH14 4AS UK