1. African Primate Initiative for Ecology and Speciation; Department of Zoology and Entomology; University of Fort Hare; Private Bag X1314 Alice 5700 South Africa
2. Ecole Pratique des Hautes Etudes; Laboratoire Paléobiodiversité et Evolution & UMR uB CNRS 6282 «Biogéosciences»; Université de Bourgogne; 6 Bld Gabriel Dijon 21000 France