1. University of StrasbourgCNRS, Immunology, Immunopathology and Therapeutic Chemistry, UPR 3572 67000 Strasbourg France
2. Graduate School of Human and Environmental StudiesKyoto University, Sakyo-ku Kyoto 606-8501 Japan
3. State Key Laboratory of Radiation Medicine and Protection, School of Radiation Medicine and Protection & School for Radiological and Interdisciplinary Sciences (RAD-X)Collaborative Innovation Center of Radiation Medicine of Jiangsu Higher Education InstitutionsSoochow University Suzhou Jiangsu 215123 China