1. Graduate School of Environmental Science Hokkaido University Sapporo 060–0810 Japan
2. Current address: Faculty of Agronomic Engineering and Forestry Zambeze University Mocuba 2403 Mozambique
3. Field Science Center for Northern Biosphere Hokkaido University Sapporo 060–0811 Japan
4. Research Faculty of Agriculture Hokkaido University Sapporo 060–8589 Japan
5. Current address: Faculty of Agro‐Food Science Niigata Agro‐Food University Tainai Niigata 959‐2702 Japan